BSc Biology, York, 1987-1990
PhD Biology, Imperial College London, 1991-1994
Research Fellow, Institute of Zoology, 1994-2000
Research Fellow, University of Cambridge, 2000-2003
Lecturer, University of Cambridge, 2003-2004
Senior Lecturer, Reader, Professor, Imperial College London, 2004-2012
Ecology and Evolution.
Linking ecological and evolutionary dynamics in theory, in the lab and in the field. I develop theory and test it using data from laboratory systems (bulb mites) and field systems (mainly mammals). Field systems include red deer in Scotland, and wolves in Yellowstone National Park.